Well, it's higher compression, combined with better torque scavenging exhaust, combined with a better flowing head, combined with a cam that breathes better with more lift and duration, combined with a better tune that throws emissions out for more power, meaning less throttle for a given tq ouput, meaning less fuel used for a given rpm/speed.
My 99 turbo Neon got over 50mpg cruising at 70mph, made 400 hp, pushed 25 psi and had a monstrous cam, 9.8:1 pistons, was over bored, had a highly ported and polished head, and 1000cc injectors. This was with 3.94 gearing, so, the shortest gears offered for the car. Probably could have gotten over 60 with the long gear tranny. When emissions are ignored power and economy gains can be rediculous.